Deforestation and reforestation are affecting biodiversity worldwide (Foley et al. 2005). The loss of forests is greatlymodifying the distribution, abundance, function and compo- sition of the biota (Ibarra and Martin 2015). In the extreme, deforestation results in species extinction (Pimm et al. 2014). In areas where human disturbed lands are abandoned (e.g. agriculture, pasture, slash and burn), the increase in natural regenerated secondary forest cover can provide habitat for many species and improve ecosystem services (Chazdon et al. 009, Poorter et al. 2016, Rozendaal et al. 2019).